| Preface | |
| To the Student | |
| Pt. I | Background to the Study of Evolution | 1 |
| Ch. 1 | Evolutionary Biology | 3 |
| Ch. 2 | A Short History of Evolutionary Biology | 17 |
| Ch. 3 | Genetics and Development | 31 |
| Ch. 4 | Ecology: The Environmental Context of Evolutionary Change | 59 |
| Pt. II | Patterns and History | 85 |
| Ch. 5 | The Tree of Life: Classification and Phylogeny | 87 |
| Ch. 6 | Evolving Lineages in the Fossil Record | 127 |
| Ch. 7 | A History of Life on Earth | 165 |
| Ch. 8 | The Geography of Evolution | 201 |
| Pt. III | Evolutionary Processes in Populations and Species | 227 |
| Ch. 9 | Variation | 231 |
| Ch. 10 | The Origin of Genetic Variation | 267 |
| Ch. 11 | Population Structure and Genetic Drift | 297 |
| Ch. 12 | Natural Selection and Adaptation | 337 |
| Ch. 13 | The Theory of Natural Selection | 365 |
| Ch. 14 | Multiple Genes and Quantitative Traits | 397 |
| Ch. 15 | Species | 477 |
| Ch. 16 | Speciation | 481 |
| Pt. IV | Character Evolution | 517 |
| Ch. 17 | Form and Function | 519 |
| Ch. 18 | The Evolution of Interactions among Species | 539 |
| Ch. 19 | The Evolution of Life Histories | 561 |
| Ch. 20 | The Evolution of Behavior | 579 |
| Ch. 21 | The Evolution of Genetic Systems | 605 |
| Ch. 22 | Molecular Evolution | 625 |
| Pt. V | Macroevolution: Evolution above the Species Level | 647 |
| Ch. 23 | Development of Evolution | 651 |
| Ch. 24 | Pattern and Process in Macroevolution | 677 |
| Ch. 25 | The Evolution of Biological Diversity | 703 |
| Ch. 26 | Human Evolution and Variation | 727 |
| Epilogue: Evolutionary Biology in the Future | 753 |
| App. I | Elementary Statistics | 755 |
| App. II | Contending with Creationism | 759 |
| Glossary | G-1 |
| Literature Cited | L-1 |
| Index | I-1 |